Anilao, Philippines Sea Slugs

The photos on these pages were taken on five trips to Anilao, Batangas, the Philippines: 3 to 18 May 2015, 6 to 18 May 2016, 5 to 19 May 2017, 5 to 15 May 2018 and 25 April to 6 May 2019. All trips were based around Club Ocellaris.

Animals were not collected and no measuring device was used. Measurements are all eyeball estimates, usually of the largest specimen we observed.

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Nudibranchia

Chromodorididae


Ardeadoris averni


Ardeadoris angustolutea

Ardeadoris cruenta

Ardeadoris egretta

Ardeadoris symmetrica

Cadlinella ornatissima

Ceratosoma gracillimum

Ceratosoma tenue

Ceratosoma trilobatum

Ceratosoma? sp. a217

Ceratosoma sp. a403

Chromodoris alcalai

Chromodoris annae

Chromodoris colemani

Chromodoris dianae

Chromodoris cf elisabethina

Chromodoris joshi

Chromodoris lochi

Chromodoris magnifica

Chromodoris strigata

Chromodoris cf strigata

Chromodoris willani

Diversidoris crocea

Diversidoris flava

Doriprismatica atromarginata

Doriprismatica paladentata

Glossodoris buko

Glossodoris sp. cf cincta

Glossodoris hikuerensis

Glossodoris rufomarginata

Glossodoris sp. a297

Goniobranchus albonares

Goniobranchus cf alius

Goniobranchus aureopurpureus

Goniobranchus coi

Goniobranchus collingwoodi

Goniobranchus fabulus

Goniobranchus fidelis

Goniobranchus geometricus

Goniobranchus hintuanensis

Goniobranchus kuniei

Goniobranchus leopardus

Goniobranchus preciosus

Goniobranchus preciosus

Goniobranchus setoensis

Goniobranchus verrieri

Goniobranchus sp. a018

Goniobranchus sp. a059

Goniobranchus sp. a285

Goniobranchus sp. a305

Goniobranchus sp. a399

Goniobranchus sp. a404

Goniobranchus sp. a529

Hypselodoris apolegma

Hypselodoris babai

Hypselodoris bollandi

Hypselodoris bullockii

Hypselodoris confetti

Hypselodoris decorata

Hypselodoris emma

Hypselodoris iba

Hypselodoris infucata

Hypselodoris kaname

Hypselodoris krakatoa

Hypselodoris lacuna

Hypselodoris maculosa

Hypselodoris cf maculosa

Hypselodoris maritima

Hypselodoris maritima
juvenile?

Hypselodoris paradisa

Hypselodoris purpureomaculosa

Hypselodoris reidi

Hypselodoris roo

Hypselodoris skyleri

Hypselodoris tryoni

Hypselodoris variobranchia

Hypselodoris whitei

Hypselodoris zephyra

Hypselodoris sp. a182

Hypselodoris sp. a224

Mexichromis mariei

Mexichromis multituberculata

Mexichromis pusilla

Mexichromis trilineata

Miamira alleni

Miamira miamirana

Miamira sinuata

Thorunna daniellae

Thorunna florens

Thorunna furtiva

Thorunna halourga

Verconia alboannulata

Verconia norba

Verconia simplex

Verconia romeri

Verconia sp. a259

Verconia sp. a481

Verconia? sp. a497

Actinocyclidae


Hallaxa cryptica


Hallaxa hileenae

Dorididae


Aldisa fragaria


Doris nucleola

Doris pecten

Discodorididae


Asteronotus cespitosus


Asteronotus hepaticus

Asteronotus mimeticus

Asteronotus? sp. a488

Atagema intecta

Atagema sp. a262

Atagema sp. a549

Avaldesia albomacula

Carminodoris bifurcata

Carminodoris estrelyado

Carminodoris flammea

Carminodoris flammea

Discodoris boholiensis

Discodoris lilacina

Discodoris cebuensis

Discodoris? sp. a440

Halgerda batangas

Halgerda dalanghita

Halgerda indotessellata

Halgerda tessellata

Halgerda willeyi

Hoplodoris rosans

Hoplodoris sp. a213

Jorunna funebris

Jorunna rubescens

Jorunna cf ramicola

Jorunna sp. a319

Jorunna sp. a107

Jorunna sp. a258

Jorunna sp. a291

Jorunna? sp. a506

Paradoris liturata

Paradoris sp. a169

Pharodoris philippinensis

Platydoris cinereobranchiata

Platydoris cruenta

Platydoris ellioti

Platydoris formosa

Platydoris inframaculata

Platydoris inornata

Platydoris cf inornata

Platydoris cf pulchra

Platydoris sanguinea

Platydoris sp. a253

Platydoris sp. a260

Platydoris? sp. a274

Platydoris sp. a407

Platydoris sp. a414

Platydoris sp. a561

Platydoris sp. a565

Rostanga? sp. a228

Rostanga sp. a490

Sclerodoris cf apiculata

Sclerodoris tuberculata

Taringa halgerda

Thordisa villosa

Thordisa sp. a063

Thordisa sp. a272

Thordisa? sp. a491

Thordisa sp. a494

discodorid sp. a094

discodorid sp. a190

discodorid sp. a300

discodorid sp. a475

Dendrodorididae


Dendrodoris albopurpura

Dendrodoris carbunculosa

Dendrodoris coronata

Dendrodoris elongata

Dendrodoris guttata

Dendrodoris krusensternii

Dendrodoris nigra

Dendrodoris cf tuberculosa

Dendrodoris sp. a068

Dendrodoris sp. a202

Phyllidiidae


Phyllidia coelestis


Phyllidia elegans

Phyllidia exquisita

Phyllidia ocellata

Phyllidia cf ocellata

Phyllidia picta

Phyllidia polkadotsa

Phyllidia varicosa

Phyllidia willani

Phyllidiella pustulosa

Phyllidiella rudmani

Phyllidiopsis annae

Phyllidiopsis burni

Phyllidiopsis cardinalis

Phyllidiopsis fissurata

Phyllidiopsis gemmata

Phyllidiopsis krempfi

Phyllidiopsis shireenae

Reticulidia fungia

Reticulidia halgerda

Hexabranchidae


Hexabranchus lacer

Gymnodorididae


Gymnodoris cf alba


Gymnodoris brunnea

Gymnodoris inornata

Gymnodoris rubropapulosa

Gymnodoris sp. a250


Gymnodoris sp. a251


Gymnodoris sp. e284

Gymnodoris sp. a353

Gymnodoris sp. a382

Gymnodoris sp. a401

Gymnodoris sp. a402

Polyceridae


Kalinga ornata

Kaloplocamus acutus

Kaloplocamus sp. a185

Kaloplocamus sp. a528

Plocamopherus maculapodium

Plocamopherus pecoso

Plocamopherus tilesii

Plocamopherus sp. a208

Plocamopherus sp. a467

Polycera abei

Nembrotha chamberlaini

Nembrotha cristata

Nembrotha kubaryana

Nembrotha lineolata

Nembrotha milleri

Nembrotha mullineri

Nembrotha sp. a376

Nembrotha sp. a465

Nembrotha sp. a558

Nembrotha sp. a566

Tambja gabrielae

Tambja morosa

Roboastra gracilis

Thecacera pacifica

Thecacera picta

Thecacera sp. a186

Thecacera sp. a173

Tyrannodoris luteolineata

Goniodorididae


Bermudella brunneomaculata

Bermudella? sp. a411

Ceratodoris kendi

Ceratodoris? sp. a295

Goniodoridella geminae

Goniodoridella sp. a356


Goniodoridella sp. 458


Murphydoris adusta

Murphydoris cobbi

Murphydoris cf maracabranchia

Naisdoris liklik

Naisdoris vitiligata

Okenia? sp. a408

Pelagella albopunctata

Pelagella longicornis

Trapania darvelli

Trapania euryeia

Trapania gibbera

Trapania scurra

Trapania squama

Trapania vitta

Aegiretidae


Aegires cf exeches


Aegires flores

Aegires gardineri

Aegires villosus

Aegires sp. a108

Aegires sp. e293

Aegires sp. a309

Aegires sp. a445

Aegires sp. a462

Flabellinidae


Coryphellina delicata

Coryphellina exoptata

Coryphellina flamma

Coryphellina lotos

Coryphellina cf lotos

Coryphellina? sp. a129

Samlidae


Samla bicolor


Samla bilas

Samla macassarana

Samla riwo

Samla? sp. a571

Babakinidae


Babakina indopacifica

Fionidae


Eubranchus putnami


Eubranchus rubropunctatus

Eubranchus sp. a225

Eubranchus sp. a396

Eubranchus sp. a572

Tenellia lugubris

Tenellia melanobranchia

Tenellia cf melanobrachia

Tenellia minor

Tenellia sp. a025

Tenellia sp. a177

Tenellia sp. a365

Tenellia sp. a405

Tenellia sp. a428


Tenellia sp. a452

Tenellia sp. a500

Tenellia sp. a501

Tenellia? sp. a515

Tenellia sp. a525

Tergiposacca longicerata

Unidentiidae


Unidentia sandramillenae

Facelinidae


Caloria indica


Caloria? sp. a113

Cratena simba

Cratena sp. a410

Cratena sp. a519

Facelina sp. a406

Facelina sp. a544

Favorinus japonicus

Favorinus mirabilis

Favorinus tsuruganus

Favorinus sp. a111

Favorinus sp. a112

Herviella claror?

Moridilla sp. a060

Noumeaella rehderi

Noumeaella sp. a417

Pruvotfolia rhodopos

Pteraeolidia semperi

Pteraeolidia semperi

Facelinid sp. a241

Facelinid sp. a327

Facelinid sp. a398

Facelinid sp. a505

Myrrhinidae


Godiva sp. a191


Godiva? sp. a350

Phyllodesmium briareum

Phyllodesmium colemani

Phyllodesmium crypticum

Phyllodesmium hyalinum

Phyllodesmium iriomotense

Phyllodesmium jacobsenae

Phyllodesmium kabiranum

Phyllodesmium karenae

Phyllodesmium koehleri

Phyllodesmium lizardense

Phyllodesmium longicirrum

Phyllodesmium macphersonae

Phyllodesmium magnum

Phyllodesmium parangatum

Phyllodesmium pinnatum

Phyllodesmium poindimiei

Phyllodesmium cf poindimiei

Phyllodesmium rudmani

Phyllodesmium tuberculatum

Phyllodesmium undulatum

Phyllodesmium sp. a043a

Phyllodesmium sp. a386

Phyllodesmium sp. a450

Aeolidiidae


Baeolidia moebii


Bulbaeolidia alba

Cerberilla affinis

Cerberilla cf albopunctata

Cerberilla sp. a153

Limenandra barnosii

Limenandra fusiformis

Arminidae


Armina cf comta


Armina cf magna

Armina major

Armina occulta

Armina scotti

Armina sp. a282

Armina sp. a378

Armina sp. a426

Armina sp. a430

Armina sp. a486

Dermatobranchus caeruleomaculatus

Dermatobranchus fortunatus

Dermatobranchus funiculus

Dermatobranchus funiculus

Dermatobranchus cf funiculus

Dermatobranchus gonatophorus

Dermatobranchus kalyptos

Dermatobranchus ornatus

Dermatobranchus phyllodes

Dermatobranchus rubidus

Dermatobranchus tuberculatus

Dermatobranchus cf tuberculatus

Dermatobranchus sp. a027

Dermatobranchus sp. a211

Dermatobranchus sp. a212

Dermatobranchus sp. a324

Dermatobranchus sp. a466

Dermatobranchus sp. a521

Doridomorphidae


Doridomorpha gardineri

Pinufiidae


Pinufius rebus

Proctonotidae


Janolus flavoanulatus

Janolus savinkini

Janolus sp. a447

Bornellidae


Bornella stellifer

Scyllaeidae


Notobryon wardi

Scyllaea fulva

Tethydidae


Melibe colemani

Melibe coralophilia

Melibe digitata

Melibe papillosa

Melibe viridis

Melibe cf viridis

Melibe sp. a052

Melibe sp. a231

Tritoniidae

Our IDs of the various Marionia species are still a work in progress. It is not clear to us how much variation might be found in some of the species. While some are easily recognized, others are difficult to distinguish from each other, and a number of those in the links below could be forms of one or a very few species.


Marionia arborescens

Marionia distincta

Marionia cf rubra

Marionia sp. a091

Marionia sp. a139

Marionia sp. a141

Marionia sp. a184

Marionia sp. a246

Marionia sp. a276

Marionia sp. a277

Marionia sp. a288

Marionia sp. a307

Marionia sp a344


Marionia sp. a385

Marionia sp. a425

Marionia sp. a455

Marionia sp. a456

Marionia sp. a551

Marionia sp. a552

Marionia sp. a554

Tritoniopsis alba

Tritonia sp. a275

Tritonia sp. a322

Tritonia sp. a345

Tritonia? sp. a448

Tritonia? sp. a555

 

Dotidae


Doto racemosa


Doto cf racemosa

Doto ussi

Doto sp. a537

Lomanotidae


Lomanotus sp. a424

Acteonoidea

Amplustridae


Micromelo guamensis

Cephalaspidea

Bullidae


Bulla ampulla


Bulla vernicosa

Haminoeidae


Aliculastrum cylindricum


Atys kuhnsi

Atys naucum

Atys sp. a303

Lamprohaminoea cymbalum

Aglajidae


Aglaja fulvipunctata


Chelidonura amoena

Chelidonura hirundinina

Chelidonura pallida

Chelidonura varians

Mariaglaja inornata

Mariaglaja tsurugensis

Noalda sp. a034

Noalda sp. 084

Noalda sp. a349

Odontoglaja guamensis

Philinopsis ctenophoraphaga

Philinopsis speciosa

Philinopsis speciosa

Philinopsis buntot

Tubulophilinopsis gardineri

Tubulophilinopsis lineolata

Tubulophilinopsis pilsbryi

Tubulophilinopsis reticulata

Tubulophilinopsis sp. a194

Tubulophilinopsis sp. a255

aglajid sp. a050

Philinidae


Philine orientalis

Gastropteridae


Gastropteron bicornutum


Sagaminopteron ornatum

Sagaminopteron psychedelicum

Sagaminopteron bilealbum

Sagaminopteron sp. a148

Sagaminopteron sp. a388

Siphopteron brunneomarginatum

Siphopteron flavolineatum

Siphopteron makasig

Siphopteron sp. a460

Diaphanidae


Colpodaspis thompsoni

Anaspidea

Aplysiidae


Aplysia nigrocincta


Aplysia oculifera

Aplysia reticulata

Bursatella leachii

Dolabella auricularia

Dolabrifera dolabrifera

Notarchus indicus

Phyllaplysia lafonti

Stylocheilus longicauda

Stylocheilus striatus

Notaspidea

Pleurobranchidae


Berthellina delicata


Euselenops luniceps

Pleurobranchaea brockii

Pleurobranchus albiguttatus

Pleurobranchus forskalii

Pleurobranchus forskalii

Pleurobranchus forskalii

Pleurobranchus grandis

Pleurobranchus grandis
juvenile

Pleurobranchus peroni

Pleurobranchus weberi

Pleurobranchus sp. a207

Tomoberthella martensi

Umbraculoidea

Umbraculidae


Umbraculum umbraculum

Sacoglossa

Cylindrobullidae


Volvatella vigourouxi

Oxynoidae


Lobiger viridis


Oxynoe viridis

Hermaeidae


Hermaea sp. a367

Limapontiidae


Costasiella kuroshimae


Costasiella usagi

Stiliger ornatus

Caliphyllidae


Cyerce elegans


Cyerce sp. a470

Polybranchia jannae

Polybranchia orientalis

Polybranchia sp. a273

Elysiidae


Elysia cf nigropunctata


Elysia marginata

Elysia pusilla

Elysia sp. a357


Elysia sp. a375

Elysia sp. a387

Elysia sp. a391

Elysia sp. a436

Elysia sp. a457

Elysia sp. e493

Plakobranchus papua

Plakobranchus sp. a567

Thuridilla albopustulosa

Thuridilla carlsoni

Thuridilla gracilis

Thuridilla undula

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Our thanks to the people at the resort Club Ocellaris: the friendly and helpful staff, always ready with meals when we returned (often late) from dives; the excellent boat operators and crews; and the terrific dive guides Peri and Alexis and others, without whom we would have seen far less. Thanks also to our fellow nudi-philic divers, all willing to share their finds so we could all see as many slugs as possible.